(+) = species; [ ssp.] = subspecies; var.= variety; syn.= synonym  

List of Butterflies of Andalusia / Lista de Mariposas de Andalucía

PAPILIONOIDEA Latreille, 1802

NYMPHALIDAE Rafinesque, 1815 ... (55 species)

Libytheinae Boisduval, 1833  (1 especie)

      Libythea Fabricius, 1807:

                (+) Libythea celtis Laicharting, 1782

Danainae Boisduval, 1833   (2 especies)

      Danaus Kluk, 1780:

                (+) Danaus plexippus L., 1758; (+) Danaus chrysippus L., 1758

Charaxinae Guenée, 1865  (1 especie)

      Charaxes Ochsenheimer, 1816:

                (+) Charaxes jasius L., 1767

Limenitidinae Behr, 1864  (1 especies)

      Limenitis Fabricius, 1807:

                (+) Limenitis reducta Staudinger, 1901   

Nymphalinae Rafinesque, 1815  (16 especies)

      Vanessa Fabricius, 1807:

                (+) Vanessa atalanta L., 1758; (+) Vanessa cardui L., 1758; (+) Vanessa virginiensis Drury, 1773

      Nymphalis Kluk, 1802:

                (+) Nymphalis polychloros L., 1758

      Aglais Dalman, 1816:

                (+) Aglais urticae L., 1758

      Polygonia Hübner, 1819:

                (+) Polygonia c-album L., 1758

      Melitaea Fabricius, 1807: (+) Melitaea didyma Esper, 1778; (+) Melitaea trivia Denis & Schiffermüller, 1775: [ssp. ignasiti Sagarra, 1926];

                (+) Melitaea cinxia L., 1758; (+) Melitaea phoebe Denis & Schiffermüller, 1775: [ssp. occitanica Staudinger, 1861];

                (+) Melitaea aetherie Hübner, 1826; (+) Melitaea parthenoides Keferstein, 1851; (+) Melitaea deione Geyer, 1832;

                (+) Melitaea celadussa Fruhstorfer, 1910 (syn. M. athalia celadussa): [var. nevadensis Oberthür, 1904, in Andalusia] [celadussa is a species: recent DNA study]

      Euphydryas Scudder, 1872:

                (+) Euphydryas desfontainii Godart, 1819: [ssp. baetica Rambur, 1858]; 

                (+) Euphydryas aurinia Rottemburg, 1775: [ssp. beckeri Herrich-Schäffer, 1851]

Heliconiinae Swainson, 1822 (8 especies)

      Argynnis Fabricius, 1807:

                (+) Argynnis paphia L., 1758; (+) Argynnis pandora Denis & Schiffermüller, 1775; (+) Argynnis aglaja L., 1758  

                (+) Argynnis niobe L., 1758: [ssp. altonevadensis Reisser, 1927]; (+) Argynnis adippe Denis & Schiffermüller, 1775 (form chlorodippe]

      Brenthis Hübner, 1819:

                (+) Brenthis daphne Denis & Schiffermüller, 1775; (+) Brenthis hecate Denis & Schiffermüller, 1775

      Issoria Hübner, 1819:

                (+) Issoria lathonia L., 1758

Satyrinae Boisduval, 1833 (26 especies)

      Lasiommata Westwood, 1841:

                (+) Lasiommata maera L., 1758; (+) Lasiommata megera L., 1767

      Pararge Hübner, 1819:

                (+) Pararge aegeria L., 1758 [ssp. aegeria]

      Coenonympha Hübner, 1819:

                (+) Coenonympha dorus Esper, 1782 [ssp. dorus, variable]; (+) Coenonympha pamphilus L., 1758: [ssp. lyllus Esper, 1806]

      Maniola Schrank, 1801:

                (+) Maniola jurtina L., 1758: [ssp. hispulla Esper, 1805]

      Pyronia Hübner, 1819:

                (+) Pyronia  tithonus L., 1771; (+) Pyronia  cecilia Vallantin, 1894; (+) Pyronia bathseba Fabricius, 1793

      Hyponephele Muschamp, 1915:

                (+) Hyponephele lycaon Kühn, 1774; (+) Hyponephele lupina Costa, 1836

      Erebia Dalman, 1816:

                (+) Erebia hispania Butler, 1868

      Melanargia Meigen, 1828:

                 (+) Melanargia occitanica Esper, 1789; (+) Melanargia ines Hoffmannsegg, 1804;  

                 (+) Melanargia lachesis Hübner, 1790; (+) Melanargia russiae Esper, 1783

      Hipparchia Fabricius, 1807:

                 (+) Hipparchia alcyone Denis & Schiffermüller, 1775;

                 (+) Hipparchia semele L., 1758; (+) Hipparchia statilinus Hufnagel, 1766; (+) Hipparchia fidia L., 1767

      Chazara Moore, 1893:

                (+) Chazara briseis L., 1764; (+) Chazara prieuri Pierret, 1837: [ssp. iberica  Oberthür, 1907).

      Pseudochazara De Lesse, 1951:

                 (+) Pseudochazara (hippolyte??) Esper, 1783 (Very probably it's a different species to hippolyte; it's necessary a molecular study, DNA/ADN):

                     [sspp. or ecological forms?: williamsi Romei, 1927; augustini Weiss, 1980; aislada Eitschberger & Steiniger, 1973; reverchoni Tarrier, 1993]

      Satyrus Latreille, 1810:

                (+) Satyrus actaea Esper, 1781

      Brintesia Fruhstorfer, 1911:

                (+) Brintesia circe Fabricius, 1775

      Arethusana De Lesse, 1951:

                (+) Arethusana arethusa Denis & Schiffermüller, 1775: [ssp./sp.? boabdil Rambur, 1839. It's necessary a study on its DNA]

LYCAENIDAE Leach, 1815 ... (46 species)

Lycaeninae Leach, 1815 (10 especies)

Neozephirus: Sabatini & Ito, 1942:

(+) Neozephirus (syn. Quercusia) quercus L., 1758: [ssp. iberica  Staundinger, 1901]

Tomares Rambur, 1840:

(+) Tomares ballus Fabricius, 1787

Callophrys Billberg, 1820:

(+) Callophrys rubi L., 1758: [ssp. fervida Staudinger, 1901]; (+) Callophrys avis Chapman, 1909

Satyrium Scudder, 1876:

(+) Satyrium spini Denis & Schiffermüller, 1775 ; (+) Satyrium ilicis Esper, 1778; (+) Satyrium esculi Hübner, 1804

Lycaena Fabricius, 1807:

(+) Lycaena phlaeas L., 1761; (+) Lycaena alciphron Rottemburg, 1775: [ssp. gordius Sulzer, 1776, var. granatensis Ribbe, 1905 in S. Andalusia]

Laeosopis Rambur, 1858:

(+) Laeosopis roboris Esper, 1794  

Polyommatinae Swainson, 1827 (36 especies)

Lampides Hübner, 1819:

(+) Lampides boeticus L., 1758

Cacyreus Butler, 1898:

(+) Cacyreus marshalli Butler, 1898

Leptotes Scudder, 1876:

(+) Leptotes pirithous L., 1758

Tarucus Moore, 1881:

(+) Tarucus theophrastus Fabricius, 1793

Zizeeria Chapman, 1910:

(+) Zizeeria knysna Trimen, 1862  

Azanus Moore, 1881:

(+) Azanus jesous Guérin-Méneville, 1849 (in Algeciras, S. Cádiz prov. Probably introduced with ornamental Acacia spp.)

Cupido Schrank, 1801:

(+) Cupido carswelli Stempffer, 1927; (+) Cupido lorquinii Herrich-Schäffer, 1847; (+) Cupido osiris Meigen, 1829

Celastrina Tutt, 1906:

(+) Celastrina argiolus L., 1758

Pseudophilotes Beuret, 1958:

(+) Pseudophilotes panoptes Hübner, 1813; (+) Pseudophilotes abencerragus Pierret, 1837

Glaucopsyche Scudder, 1872:

(+) Glaucopsyche alexis Poda, 1761; (+) Glaucopsyche melanops Boisduval, 1828: [ssp. algirica Heyne, 1895]

Iolana Bethune-Baker, 1914:

(+) Iolana iolas Schultz, 1905: [ ssp. saritae Fernández-Rubio, 1973]

Kretania Beuret, 1959:          (Note: according to a recent molecular study, the "pylaon" group is included in the genus Kretania ]

(+) 'Plebejus' pylaon Fischer von Waldheim, 1832 [ssp. hespericus Rambur, 1839 [final taxonomic status pending molecular study, DNA/ADN]

Plebejus Kluk, 1802:

(+) Plebejus argus L., 1758:  [ssp. hypochionus Rambur, 1858]; (+) Plebejus idas L., 1758: [ssp. nevadensis Oberthür, 1896]   

Polyommatus Latreille, 1804:

(+) Polyommatus thersites Cantener, 1835;

(+) Polyommatus icarus Rottemburg, 1775 [var./hrybrid/form? abdon Aistleiner & Aistleiner, 1994: pending of a molecular study, DNA];

(+) Polyommatus celina Austaut, 1879 [syn. P. icarus celina]: Its DNA is very different to P. icarus, genetic divergence > 5% (distribution: see);

(+) Polyommatus (Plebicula) golgus Hübner, 1813; (+) Polyommatus (Plebicula) sagratrox Aistleitner, 1986: pending of a molecular study, DNA; 

(+) Polyommatus (Plebicula) nivescens Keferstein, 1851; (+) Polyommatus ( ) amanda Schneider, 1792: [spp. tova  Higgins, 1948];

(+) Polyommatus ( ) escheri Hübner, 1823; (+) Polyommatus (Lysandra) bellargus Rottemburg, 1775;

(+) Polyommatus (Lysandra) albicans Gerhard, 1851; (+) Polyommatus (Lysandra.) hispana H.-Schäffer, 1852 [in Andalusia?]

Agrodiaetus Hübner, 1822                               (Note: Agrodiaetus is regarded a subgenus of Polyommatus by some recent authors)

(+) Agrodiaetus violetae Gómez-Bustillo, Expósito & Martínez, 1979: [ssp. violetae; ssp. subbaeticus Gil-T. & Gil-Uceda, 2005 (syn. A. fabressei subbaeticus)]

Cyaniris Dalman, 1816:

(+) Cyaniris semiargus Rottemburg, 1775: [ssp. tartessus Gil-T. & Huertas, 2007]

Agriades Hübner, 1819:

(+) Agriades zullichi Hemming, 1933

Eumedonia Esper, 1780:      (Note: Eumedonia  is a different genus to Aricia  according to recent molecular studies on their DNA)

(+) Eumedonia eumedon Esper, 1780: [ssp. rondensis Gil-T. & Moreno-Benítez, 2012; ssp. axarquia Gil-T., 2004; ssp. mariensis Eitschberger & Steiniger,1975]

Aricia Reichenbach, 1817:

(+) Aricia morronensis Ribbe, 1910: [ssp. morronensis; ssp. ramburi Verity, 1929]

(+) Aricia cramera Eschscholtz, 1821: [very different genetically of A. agestis]

(+) Aricia artaxerxes Fabricius, 1793: [ssp. montensis Verity, 1928; -different species or subspecies of artaxerxes according to several authors]

PIERIDAE Swainson, 1820 ... (19 species)

Dismorphiinae Schatz, 1886 (1 especies)

Leptidea Billberg, 1820:

(+) Leptidea sinapis L., 1758

Coliadinae Swainson, 1821 (4 especies)

Colias Fabricius, 1807:

(+) Colias alfacariensis Ribbe, 1905 (syn. C. australis Verity); (+) Colias crocea Geoffroy, 1785

Gonepteryx Leach, 1815:

(+) Gonepteryx rhamni L., 1758; (+) Gonepteryx cleopatra L., 1767

Pierinae Swainson, 1820 (14 especies)

Anthocharis Boisduval, Rambur & Graslin, 1833:

(+) Anthocharis cardamines L., 1758; (+) Anthocharis belia L. 1767: [ssp. euphenoides Staudinger, 1869]

Zegris Boisduval, 1836:

(+) Zegris eupheme Esper, 1804: [ssp. meridionalis Lederer, 1852]    

Euchloe Hübner, 1819:

(+) Euchloe tagis Hübner, 1804 (ssp. tagis = a cline in the Iberian peninsula, varities: gradual change in characters (markings or colour)]

(+) Euchloe crameri Butler, 1869; (+) Euchloe belemia Esper, 1800 [ssp. belemia];

(+) Euchloe bazae Fabiano, 1993 (syn. E. charlonia bazae] 

Aporia Hübner, 1819:

(+) Aporia crataegi L., 1758

Pieris Schrank, 1801:

(+) Pieris brassicae L., 1758; (+) Pieris rapae L., 1758; (+) Pieris mannii Mayer, 1851; (+) Pieris napi L., 1758

Pontia Fabricius, 1807:

(+) Pontia daplidice L., 1758

Colotis Hübner, 1819:

(+) Colotis evagore Klug, 1829: [ssp. nouna Lucas, 1949]

PAPILIONIDAE Latreille, 1802 ... (4 species)

Parnassiinae Duponchel, 1835 (1 especies)   

Parnassius Latreille, 1804:

(+) Parnassius apollo L., 1758: [ssp. nevadensis Oberthür, 1891 ...]

Zerynthiinae Grote, 1899 (1 especie) (Note: currently Zerynthiinae is included in Parnassiinae)  

Zerynthia Ochsenheimer, 1816:

(+) Zerynthia rumina L., 1758: ssp. rumina (a cline in Spain: gradual change in characters (markings or colour)]

Papilioninae Latreille, 1802  (2 especies)   

Papilio Linnaeus, 1758:

(+) Papilio machaon Linnaeus, 1758

Iphiclides Hübner, 1819:

(+) Iphiclides podalirius Linnaeus, 1758: [ ssp. feisthamelii Duponchel, 1832]

HESPERIOIDEA Latreille, 1809

HESPERIIDAE Latreille, 1809 ... (22 species)

Pyrginae Burmeister, 1878  (16 especies)   

      Erynnis Schrank, 1801:

(+) Erynnis tages L., 1758

      Carcharodus Hübner, 1819:

(+) Carcharodus alceae Esper, 1780; (+) Carcharodus tripolinus Verity, 1925; (+) Carcharodus lavatherae Esper, 1783;

(+) Carcharodus baeticus Rambur, 1839; (+) Carcharodus flocciferus Zeller, 1847

      Spialia Swinhoe, 1912:

(+) Spialia sertorius Hoffmannsegg, 1804

      Muschampia Tutt, 1906:

(+) Muschampia proto Ochsenheimer, 1808

      Pyrgus Hübner, 1819:

(+) Pyrgus malvoides Elwes & Edwards, 1897 (syn. P. malvae malvoides); (+) Pyrgus armoricanus Oberthür, 1910;

(+) Pyrgus alveus Hübner, 1803; (+) Pyrgus serratulae Rambur, 1839; (+) Pyrgus onopordi Rambur, 1839;

(+) Pyrgus cirsii Rambur, 1839; (+) Pyrgus carthami Hübner, 1813 (syn. P. fritillarius)

Hesperiinae Latreille, 1809  (7 especies)

      Thymelicus Hübner, 1819:

(+) Thymelicus lineola Ochsenheimer, 1808; (+) Thymelicus sylvestris Poda, 1761; (+) Thymelicus acteon Rottemburg, 1775

      Hesperia Fabricius, 1793:

(+) Hesperia comma L., 1758

      Ochlodes Scudder, 1872:

(+) Ochlodes sylvanus Esper, 1778 (syn. Ochlodes venata faunus Turati 1905)

                                    Taxonomic note: O. sylvanus sometimes called O. faunus (syn. O. venata faunus) occurs in Europe and Asia east to China;

                                    O. venata from China east to Korea and Japan. They are possibly sympatric in China]  

      Borbo Evans, 1949:

(+) Borbo borbonica Boisduval, 1833: [ssp. zelleri Lederer, 1855]

      Gegenes Hübner, 1819:

(+) Gegenes nostrodamus Fabricius, 1793

List of Butterflies of the Canary islands / Lista de Mariposas de las islas Canarias

PAPILIONOIDEA Latreille, 1802

NYMPHALIDAE Rafinesque, 1815 ... (12 species)

Danainae Boisduval, 1833  (2 species)

     Danaus Kluk, 1780:

            (+) Danaus plexippus L., 1758; (+) Danaus chrysippus L., 1758  

Nymphalinae Rafinesque, 1815  (5 species)

      Vanessa Fabricius, 1807:

            (+) Vanessa atalanta L., 1758; (+) Vanessa vulcania Godart, 1819 (syn. V. indica vulcania);

            (+) Vanessa cardui L., 1758; (+) Vanessa virginiensis Drury, 1773

     Hypolimnas Hübner, 1819:

            (+) Hypolimnas misippus L., 1764 (migrant)

Heliconiinae Swainson, 1822 (2 species)

      Argynnis Fabricius, 1807:

            (+) Argynnis pandora Denis & Schiffermüller, 1775: (ssp. seitzi Fruhstorfer, 1908]

      Issoria Hübner, 1819:

            (+) Issoria lathonia L., 1758

Satyrinae Boisduval, 1833 (3 species)

      Pararge Hübner, 1819:

            (+) Pararge xiphioides Staudinger, 1871

      Maniola Schrank, 1801:

            (+) Maniola jurtina L., 1758: [ssp. jurtina; var. fortunata Alpheraky, 1889]

      Hipparchia Fabricius, 1807:

            (+) Hipparchia wyssii Christ, 1889:          [I believe that some of the sspp. described are only varieties]

                      [ssp. wyssii (Tenerife island); ssp./var.? tamadabae Owen & Smith, 1992 (Gran Canaria); ssp. tilosi Manil, 1984 (La Palma);  

                       ssp./var.? gomera Higgins, 1967 (La Gomera); ssp. bacchus Higgins, 1967 (El Hierro)]

                      (some differences mentioned in preimaginal stages, mainly in the egg, but would require a molecular study (pending) may determine its final taxonomic status)

LYCAENIDAE Leach, 1815 ... (9 species)

Lycaeninae Leach, 1815

      Lycaena Fabricius, 1807:

            (+) Lycaena phlaeas L., 1761

Polyommatinae Swainson, 1827 (8 species)

      Lampides Hübner, 1819:

            (+) Lampides boeticus L., 1758

      Cacyreus Butler, 1898:

            (+) Cacyreus marshalli Butler, 1898

      Leptotes Scudder, 1876:

            (+) Leptotes pirithous L., 1758

      Zizeeria Chapman, 1910:

            (+) Zizeeria knysna Trimen, 1862

      Polyommatus Latreille, 1804:

            (+) Polyommatus celina Austaut, 1879 [syn. P. icarus celina]: Its DNA is very different to P. icarus, genetic divergence > 5%.

      Aricia Reichenbach, 1817:

            (+) Aricia cramera Eschscholtz, 1821

     Azanus Moore, 1881

            (+) Azanus ubaldus Cramer, 1782 (in S. Gran Canaria island. Probably introduced with ornamental Acacia spp.)

     Cyclyrius Butler, 1897:

            (+) Cyclyrius webbianus Brullé, 1839

PIERIDAE Swainson, 1820 ... (8 species)

Coliadinae Swainson, 1821 (3 species)

      Colias Fabricius, 1807:

             (+) Colias crocea Geoffroy, 1785  

      Gonepteryx Leach, 1815:

            (+) Gonepteryx cleobule Hübner, 1825:

                      [ssp. cleobule (Tenerife); ssp. eversi  Rehnelt, 1974 (La Gomera); ssp. palmae Stamm, 1963 (La Palma)]

                [The genetic distance between these taxa (around 2%), mentioned by some authors, it is not sufficient reason to consider these taxa as species]

      Catopsilia Hübner, 1819

                (+) Catopsilia florella Fabricius, 1775

Pierinae Swainson, 1820 (5 species)

      Euchloe Hübner, 1819:

            (+) Euchloe belemia Esper, 1800:                  [Note: I consider these taxa as subspecies according to the results of Suárez et al. (2009)]  

                      [ssp. hesperidum Rothschild, 1913 (Fuerteventura); ssp. eversi Stamm, 1963 (Tenerife);

                       ssp. grancanariensis Acosta, 2008 (Syn. constantini Back) (Gran Canaria)]

            (+) Euchloe charlonia Donzel, 1842: [ssp. charlonia]

      Pieris Schrank, 1801:

            (+) Pieris cheiranthi Hübner, 1808: [ssp. cheiranthii (Tenerife); ssp. benchoavensis Pinker, 1968 (La Palma)];

            (+) Pieris rapae L., 1758

      Pontia Fabricius, 1807:

            (+) Pontia daplidice L., 1758

HESPERIOIDEA Latreille, 1809

HESPERIIDAE Latreille, 1809 ... (1 species)